Teton Shrubs – Spring

The Valley is full of flowers in June.  So many so that here we present them by type—here are the shrubs, which come into their own when in full flower.  Separately, we will post Wildflowers in Sageflats and Hills/Sun and Wildflowers in Forests/shade.

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Most shrubs are most appreciated in spring when they bloom, although some will have colorful foliage and fruits for a short time in fall.  The rest of the year, shrubs are usually overlooked as just green bushes or just twigs.  So now is the time to celebrate shrubs.

Note: Shrubs are woody plants usually between 6”-20’ tall with multiple stems vs. 1 or 2 trunks of a tree.

The following are more or less in flower sequence within their group.

First to Start:

WillowsSalix sp. –  There are many, many types of willows of all sizes. They are very important for wildlife: pollinating bees, nesting birds, browsing moose, and dam-building beaver to name a few.

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We usually welcome their early spring catkins—pussy willows.  They come in male and female versions, are pollinated by insects (not by wind, as previously thought) starting in April.  By late June the female capsules are bursting with thousands of seeds attached to a tufts of fluff being dispersed by wind.  Along with their relatives the cottonwoods, they are creating blizzards.

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The teensie seeds are viable for only a few days, and they must land on moist open ground to germinate. Only a couple of seeds out of a million will sprout and grow up at all. Plants, though, can propagate vegetatively from broken stems stuck in the mud, as along flooded rivers or around beaver ponds.

Utah HoneysuckleLonicera utahensis – has oval leaves that are arranged opposite each other on the stems.  The trumpet-like, pale yellow flowers are found in pairs, their ovaries attached at the very base.  Later twin red berries will loll upon the green of the leaves…but not until late July.

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Oregon GrapeMahonia repens – is one of our few evergreen shrubs and is particularly tough.  It grows in sun or shade, often on very rocky soils.  Several thick holly-like leaflets form along the central petiole to comprise a compound leaf. In spring the “evergreen” leaves remain, although a bit tattered, until the new leaves replace them.  The flowers are remarkably fragrant and last a long time.  The underlying bark is yellow with “berberine” which has medicinal properties.

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Mountain Maple  – Acer glabrum – has inconspicuous flowers that are pollinated by wind or perhaps some insects?  Each flower is usually male or female to prevent self-pollination.  By now in late June you can see the beginning of the fruits forming – the double winged samaras – that will twirl to new realms later in the fall.

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The leaves are 3-5 lobed, with some saw-like teeth along the edges.  Often I see the leaves mottled with a deep red “fungus”.  The velvet red splotches are actually formed by tiny galls created by mites:  https://www.canr.msu.edu/news/velvet_galls_caused_by_tiny_eriophyid_mites

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A June Parade of White Blooms in the Rose Family:

These common shrubs grow from a few feet to up to almost 20’ tall.  They all have five green sepals, five white petals, and anthers (the pollen bearing parts) in multiples of 5. These three whorls merge into a cup-like hypanthium that will later swell and protect the seeds. Hypanthium structures are common in the Rose Family. In the very center of this cup-like feature, sit the female ovaries and eggs that will form the seeds.  The fruits will be a combination of swollen protective hypanthiums and the mature seeds in the center. (image wikipedia)

Hypanthium - Wikipedia

All these plants are related to cultivated fruits we eat: apples, plums, pears, peaches—also in the Rose Family.  Our native roses blooms in July–very soon! and produce “hips” – the tough hypanthium with dry fruits/seeds inside.RosaSp_Fr_dis_CU_NER_11.11.18_1b_Q1_crp_5x3_200

More or less in order of bloom:

Serviceberry/SaskatoonAmelanchier alnifolia – has oval 1-1.5” leaves with a few teeth arrayed around the upper ½.  Flowers are arranged along a central stalk.  Being one of the earliest to bloom, it will also be the earliest to fruit.

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ChokecherryPrunus virginiana – has 4-5” oblong, pointed leaves with fine teeth all along the edges.  The dozens of flowers are stiffly arranged on the long central stalk.  All the parts of the plant have a chemicals that can produce poisonous cyanide under certain conditions.  For instance, if cattle eat too many of the spring leaves they can become very sick.  Amateur entomologists have used the leaves and bark in jars to kill insect specimens.  On the other hand, western tent caterpillars thrive in their branches, and birds and other mammals will eat the fruits come fall (the particularly toxic pits pass on through).

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Black HawthornCrataegus douglasii – are easy to ID with their ½-1” thorns.  The broad oval leaves are coarsely toothed.  The flowers are in branching bunches or cymes.  In August, the Moose-Wilson Road is often closed due to bears feasting on the berries.

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Mountain Ash/Greene’s AshSorbus scoparia – begins to bloom in late June in slightly cooler locations.  The leaves are “compound” with many leaflets coming off a central stalk.  The stalk then attaches to the twig.  The tiny flowers form a great puff of creamy white flowers at the end of the stems.

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NinebarkPhysocarpus malvaceus – is found in the southern part of Jackson Hole flowering at the end of June. Unlike other white bloomers above, it will have dry fruits.  Leaves are slightly lobed and toothed.

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Yellow Members of the Rose Family Join In the Parade of Bloom:

Not all members of the very large Rose Family make luscious fruits.  While also having hypanthiums, the cups remain thin and brown cradling dry seed-like fruits (achenes) inside. For now enjoy the flowers.

Antelopebrush/BitterbrushPurshia tridentata – is especially abundant this year.  You can smell the sweet yellow flowers before you come over the rise of an open hillside.  Plants are often intermixed with sagebrush, growing about the same size.

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The nutritious plants are valued wildlife browse throughout the year, but particularly in late fall and winter by moose.  Ants and mice relish the seeds.  Also these plants can fix their own nitrogen with the help of bacteria growing in the roots.  Therefore, these shrubs can grow in many tough conditions.  Note their small leaves are three-tipped like those of sagebrush, but they are greener overall, and the edges curl under.

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Shrubby Cinquefoil –  Potentilla/Diasphora fucticosa/Pentaplylloides floribunda – is a favored landscape plant for its yellow flowers in early to mid-summer and its low maintenance.  It can grow in a variety of soils from sun to shade – but prefers a bit of moisture, especially when in sun.

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Nurseries sell a range “cultivars” with white to deep orange flowers.  Our wild native is yellow.

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More June Shrubs of Various Sorts –

These four shrubs prefer more or less moist and/or cool areas and grow into substantial plants.

SilverberryEleagnus commutata – has been flowering under the cottonwoods along the Snake and Gros Ventre Rivers from Wilson north. They are easily seen from the dikes and from Fall Creek Road.

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The 2-3” silvery oblong leaves stand out in the shade on 6’-tall, erect colonial plants.  Hidden in the foliage are yellowish, highly fragrant tubular flowers.

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Red ElderberrySambucus racemosa – looks at first glance like Mountain Ash with its white bunches of small flowers and compound leaves.  However, it is overall heftier with thicker stems and heavier leaves, and notably the compound leaves are opposite each other, not alternate up the stems.   Also, plants have a foul odor if pinched or crushed.  Its European relative – Sambucus nigra – is the source of sambucol, the anti-viral flu mediation.

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TwinberryLonicera involucrata – is also a substantial shrub usually found near water. The 3-6” oval leaves are opposite on the stem, and the flowers are formed in pairs. (It is a relative of the early flowering relative Utah honeysuckle – see above.) Flowers are mostly yellowish and hairy with a “bract” at their base that can be greenish yellow but later turns deep maroon.  LoniInvo_FlCU_Lvs_7.6.16_2_Q1_5x3_180_blgKeep an eye out for caterpillars of Gillette’s Checkerspot butterfly (photo credit: Wikipedia) which require this species for their host.

Euphydryas gillettii - Wikipedia

Red-stemmed DogwoodCornus stolonifera – is loved by moose, other wildlife, and landscapers.  In fact it is an appealing “ornamental” plant for its tidy oval leaves set opposite each other on the red stems, bunches of white flowers,  and later white berries.  Red winter twigs are a cheerful contrast against snow. “Cultivars” have been selected with brighter red stems, variegated leaves, bigger flowers and berries.  The moose relish it as is, and the birds will carry off the fruits when ready. A great plant for a “wildlife friendly” garden.

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Unnotables:

Junipers are evergreen conifers.  The tree-like Western Juniper –  Juniperus scopulorum – grows in all shapes and sizes on our dry buttes.

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This species has scale-like leaves.

JuniScop_lvfrCU_GameCrk_9.20.19_1a_Q1_5x3_180 The sprawling Common JuniperJ. communis – has sharp needles that are in whorls of 3 and grows here and there in sunny spots.  JuniComm_stfr_MWRdN_11.29.1_Q1_crp_5x3_180_blgBoth produce “berries” which are technically fleshy cones that are relished by Townsend’s Solitaires and Cedar Waxwings.  However, few realize that the cones take 18 months to form, starting in June, and only on “female” plants.

Not to get into the complications of “naked seed” plants and their evolution, suffice it to say that pollen is released from tiny structures,

JuniScop_flM_GameCrk_6.10.20_3_Q2_crp_5x5_180_blgand with luck land on females cones of a different plant.  You need a microscope to really see what is going on, but with a handlens, you can observe the first bulge of reproduction.

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Others shrubs not mentioned: The gooseberries/currants – Ribes sp., various huckleberry relatives – Vaccinium sp., and a few inconspicuous wind pollinated species.  We save those for later.

Enjoy the shrubs of spring…more to come this summer!

Frances Clark, Program Coordinator

Wilson, WY

June 26, 2020

Spring Flowers Mid-May 2020

As the valley greens up and we need a respite from the world of Covid-19, many of us are out and about looking for flowers.  DryKnoll_NendNER_5.14.20_1_Q2_fix_5x3_180As I write, some of our regular haunts in Grand Teton National Park are still closed (but about to open!).  However, there are many other places for botanical forays.

Bridger-Teton National Forest has various accessible habitats such as more south-facing grassy slopes mixed with sagebrush and dry rocky slopes where the snow melts early.   Josie’s Ridge, Cache Creek, and the lower slopes of Munger Mountain are all good jaunts.  Dry hills to the north and east of Flat Creek Road; Crystal Butte near Jackson; Game Creek south of town; and Poison Creek southeast of Hoback are other accessible points.  Please always respect boundaries and watch your step—many of the plants are small and fragile.

Many common flowers are presented here.   Go to past posts of “What’s in Bloom?” for additional and different details.

Earliest flowers found to the south of Jackson a month ago are now fading but are still visible farther north: 

Turkey PeasOrogenia linearifolia – have the tiniest of white flowers and skimpy linear leaves.  The cluster if often smaller than the size of a penny.OrogLin_FlSt_Pen_WilBrk_41012_1bsm

Two early look-alike buttercups perk up the dried grasses:  Sagebrush Buttercups – Ranunculus glaberrimus –  have simple leaves—although when they stretch out they can have two lobes. 

Utah Buttercups leaves are 3-lobed from first emergence.  RAnJov_flCU_MurCtr_41112_1b_fix_5x3_180

Steer’s-headsDicentra uniflora – are easily overlooked until you get the search image of the leaves…then look for the  flowers which have an obvious western motif. DiceUnif_fllf_CrystBut_5.5.20_1_Q2_5x3_180

Slightly later or in different locations Spring BeautiesClaytonia lanceolata – are sprinkled about in light shade or sun where snow has just melted. Leaves are opposite, and the flowers are pinkish with delicate, pinker veins that lead pollinators to the center of the flower.ClayLanc_flhab_MuriRch_4.21.18_1_crp5x3_180fix

Ever-popular YellowbellsFritillaria pudica – grace hillsides and sageflats with their charming bells.FriPudi_fl_SchwLd_bench_fl_52011_1a_5x3_180fix

As we move into mid-May, species found on grassy slopes and amidst sage brush include:

Yellow violets – Viola nuttalii varieties – have a range of leaf shapes and sizes which confuse exact identification to variety.    ViolNuttVallcf_flhab_FltCRkHills_5.8.20_2b_Q2_crp_5x3_180Goosefoot VioletViola purpurea – is easy to ID because of its distinctive webbed-foot leaves.ViolPurpVen_HabFl_DMansB_52113_1_5x3_180

Oblongleaf Bluebell – Mertensia obtusifolia – flowers dangle, changing color from pink to blue when ready to attract pollinators. MertOblo_habfl_GameCrk_5.9.20_1_Q2_5x3_180

Shooting StarDodocatheon conjugens – folds back its petals and has little knobs for bees to cling to—it is “buzz” pollinated. A bee lands, hangs, vibrates its wing muscles and  then pollen grains sift out of the purple anthers and land on the bee’s belly.  The bee flies to another flower with a stigma sticking out which then tags the pollen. DodoConj_flhab_CrystBut_5.5.20_2_Q2_crp_3x4.5_180

Nuttall LarkspurDelphinium nuttallianum – will soon become more abundant and attract hummingbirds.DelpNutt_fl_SkiLakeTr_73111_3aQ1Sm

Dry slopes are some of my favorite spring botany spots:BTNF_FltCRkHills_vw_spring_5.8.20_4a_fix_%x3_180

Wonderfully fragrant Hood’s PhloxPhlox hoodii – has tight needle-like leaves with cobwebby hairs that help distinguish it from the later blooming Multiflora Phlox.  Flowers can range from light to dark blue on different plants of the same species.PhloHood_habfl_CU_NendNER_5.14.20_1_5x3_180_Q2

Low pussytoesAntennaria dimoprha – are often overlooked even when in flower – its tiny composite flowers are all male or female on separate plants.  Flowers are wind pollinated or females can produce seeds without a male around.AnteDimo_flCU_CrystBut_5.5.20_1a_Q1_5x3_180

Pursh’s milkvetchAstragalus purshii – has typical pea-like flowers emerging beyond a clump of compound ladder-like silvery leaves.  The flowers are creamy white. The genus of the Pea Family can be IDed, in part, by the two “wing” petals being longer than the “keel”, which here is purple tipped.AstrPurs_flhab_rock_CrystBut_5.5.20_1_Q2_crp_5x3_180

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Bessey’s LocoweedOxytropus besseyi – has elegant fists of flowers above similar leaves to milkvetch.  The pea-like are very similar (except for its color!); however, the “wing” petals are shorter than the pointy “keel” in this genus of the Pea Family.OxytBess_flhab_NendNER_5.14.20_1a_Q2_53_180

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Whitlow GrassesDraba sp.- were abundant on a very dry slope.  ID of this confusing genus of mustards requires the fruits to ripen.  Hairs also help!DrabSp_fhab_rock_NendNER_5.14.20_1_Q2_5x3_180

Mountain RockcressBoechera sp. – is another Mustard Family member awaiting full ID.  All mustards have 4 sepals, 4 petals, 6 stamens – 2 shorter than the other 4, and a single pistil.  The pistils of the family grow into fruits of various shapes—the key ID feature needed. (photo by Susan Marsh)BoecSp_FlCU_rockcress_by SM5.14.20_crp_5x5_180

Desert PaintbrushCastilleja chromosa – is the first of several paintbrushes that will bloom over the next several months.  Its orange-red, day-glow flowers decorate dry hillsides amidst colorful rocks.CastChrom_habfl_FltCRkHills_5.8.20_1_Q2_5x3_180

Below are some of the plants, which for some reason, have been given less prominence in past posts—not sure why:

PasqueflowersAnemone patens – are an early spring favorite for obvious reasons!AnemPate_flhab_CrystBut_5.5.20_1a_Q2_5x3_180

Diamondleaf SaxifrageSaxifraga rhomboidea – is often overlooked.  The basal rosette of rhomboid leaves gives rise to a single 2-9” stalk with a cluster of several white flowers. SaxiRhom_budlvs_FltCRkHills_5.8.20_2_Q1_crp_5x3_180SaxiRhom_flCU_ant_JosieRidge_52311_2_crp_3x3_180

Oregon GrapeMahonia repens – is common in many habitats, but we often forget to get down and sniff the wonderfully fragrant flowers!  But watch your nose on the spiny, tough evergreen leaves.MahoRep_fl_DinNM_513112_2_5x3_180

Wyoming KittentailsBesseya wyomingensis – have no showy petals.  The color comes from the purple-blue stamens that unfold to yield white pollen.BessWyom_habfl_NendNER_5.14.20_3a_Q2_crp_3x4.5_180

Best to get out now to see these early spring flowers before you are distracted by so many more flowers to come, such as Balsamroot!BalsSagi_habfl_FltCRkHills_5.8.20_1_fix_5x3_180Happy Spring.

Frances Clark, Teton Plants Program Coordinator

P.S. As always, we appreciate any corrections, suggestions, or other comments!

Wildflower Crescendo at High Elevations

On Thursday, August 22, a friend and I took the JH Mountain Resort tram up to 10,400’ elevation for a hike down through Cody Bowl along the Rock Springs Loop trail*.

While at first glance the slopes appear barren,Rendv_top_vwN_8.22.19_1_5x3_180

we were delighted to find so many wildflowers – from early summer favorites to late season popups – blooming together. 

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This spring snows were deep and melted late on slopes and basins in the irregular and spectacular topography of the mountains.  Consequently, there was an unexpected crescendo of bloom  at the end of a summer-long symphony of flowers. 

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While we took the tram up and hiked only a few miles, others of you may be able to hike farther and higher to find additional locations still colorful with wildflowers.

Below is a showcase of flowers on Rendezvous Mountain with habitat and ID tidbits.  Avid or novice wildflower watcher, please enjoy!

The very top of the ridge in loose rocks and drying winds

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A few Old-Man-of-the-MountainHymenoxys grandiflora – still bloom among the gravelly soils. They indeed look like wizened sages of the alpine.

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Mountain Dryad – Dryas octopetala – a member of the Rose Family which can fix its own nitrogen, inhabits these lean, exposed soils, growing in very low mats.  After its saucer-like yellow flowers are pollinated, they produce twisted clusters of fruits

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which will fly off individually to find a place to germinate in time.

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Gordon IvesiaIvesia gordonii – has tight heads of several yellow flowers that are beginning to fade.  The “pinnately-divided” leaves are at the base: the tiny, hairy leaflets branch off from the center of the leaf. This is also in the rose family.

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D—! Yellow Composites:

Two spreading yellow composites can form large patches of yellow:

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Shrubby Goldenweed – Haplopappus suffructicosus – is finishing bloom along the mountain shoulder. The whole plant is glandular and fragrant.  Wide-spreading flowers are at the tips of 1-2’ woody stems. The oblong 1-2” leaves alternate up the stem.  

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Longleaf ArnicaArnica longifolia – is just beginning to flower near the summit.  The largest of of our arnicas, these 2-2.5’ tall stems have 5-10 pairs of long opposite leaves, and several relatively small yellow flower heads at the top.

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Largeleaf Arnicas form extensive colonies in rocky seeps and wet talus visible from the tram.

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Two-leaf GroundselPackera/Senecio dimorphophylla – is a high-elevation species with succulent leaves, including at least one relatively large, slightly lobed leaf clasping the stem.  The “two-leaf” of the name indicates the great variety in leaf shapes that adds to ID woes.

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It can be mistaken for another mostly lower-elevation Twisted-leaf groundselPackera/Senecio streptanthifolia. It has thinner leaves, longer stems to the flowers, and is generally less compact.  But they are confusing!

Several species familiar from lower elevations:

 

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Common YarrowAchilea millefolium – is one of the most adaptable and widespread species growing not only within a full range of elevations, but also it spreads around the Northern Hemisphere.

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False or Mountain DandelionAgoseris glauca var. dasyphyllum – grows to about 6-8” with large heads of ray flowers. This native is commonly seen in the sageflats in May.

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Rocky Mountain GoldenrodSolidago multiflora – grows from 4” to 18” high depending on elevation and associated conditions. The tiny flowerheads have about 10-20 ray flowers and the lower leaves have “ciliate” hairs on the petioles.

Three Fleabane Daisies and Asters look very similar. They tend to grow 1-2.5’ high with many oval leaves alternating the stem. They have blue flowers.  Turn over the flower-heads and look at the “bracts” that form the green protective structure around the base.  They are different!

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Fleabane DaisiesErigeron sp. – have narrow, equal-length bracts. 

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Peregrine DaisyE. peregrinus has wider ray flowers (they look like petals)

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than Oregon DaisyErigeron speciosus (above).

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Leafybracted AsterSymphiotrichum foliaceum – has wide ray flowers (like Peregrine Daisy) but has broad leaf-like bracts. The low growing alpine variety is “apricus” if you are into the details.

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This slope is covered with phlox-like Nuttall’s LeptosiphonLeptosiphon/Linanthus nuttalii – with dashes of common Lewis’ FlaxLinum lewisii – which are also found at lower elevations.

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On the same slope is a mix of blue flowers: flax and penstemon.

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Thin-stem PenstemonPenstemon attenuatus – seems to be one of the most common of the confusing beard-tongues.  I look for the glandular hairs on the flowers, acute sepals, and inside the smooth anthers spreading at 180 degree angle to help with ID.   In any case penstemons have opposite leaves, blue “irregular” flowers with curled up stamens along with a “beard’s-tongue” inside.

These subalpine species grow in patches with sufficient moisture and nutrients :

 

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Death CamusZigadenus elegans – has beautiful whitish green flowers with heart-shaped yellow pools of nectar on the the 6 petals (technically here called tepals).

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It has 6 stamens that stand around the center pistil which has 3 stigmas.  Flower parts in multiples of 3 is a good indication that it is in the Lily Family.  Compare it to your Easter Lily.

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Coil-Beak LousewortPedicularis contorta – has 1-foot+ stalks arrayed with white flowers that twist around their bee pollinators to position them for prescise fertilization. The lower leaves are pinnately divided, helping to differentiate it from the similar Curled Lousewort – P. racemosa – whose leaves are only toothed. 

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Here Coil-beak Lousewort grows with Sulphur Paintbrush – Castilleja sulphurea

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Bog/Explorer’s GentianGentiana calycosa – greets you with blue goblets of flowers.

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The interior lines and spots draw pollinators deep inside searching for nectar which is at the very bottom.  As a consequence, the pollinator rubs against the male anthers or female stigma (they are receptive at different times) effecting fertilization for new seeds to come.

 

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Parry’s CatchflySilene parryi – has a swollen calyx decorated with elegant lines: They glow like Chinese lanterns. This alpine native is related to the common, weedy bladder campion – Silene latifolia – that pops up in our gardens.

Tucked into talus rock of Cody Bowl:

 

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Moss CampionSilene acaulis – is often the first alpine to bloom. This tightly growing, mounded “cushion plant” is still blooming and is also forming fruits. Plants a few inches across and an inch high can be dozens of years old. This is related to the much larger Parry’s Catchfly (see above).  Both are related to carnations!

Two miniature 2-4”-tall aster-like plants are readily confused (by me anyway!). They have single blue flowerheads at the top of short stems. The leaves are mostly clustered at the base:

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Alpine AsterOreostema alpigenus – has deep-blue flowers and long leaves. Note the long tap root that has been unearthed.  These are very old plants.

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Alpine TownsendiaTownsendia montana – has much shorter, spoon-shaped leaves.

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American ThorowaxBupleurum americanum  – is a member of the Parsley Family with an umbrella-like arrangement of flowers and fruits.

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This species is unusual in having undivided leaves. It is common among the rocks here.

 

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Tweedy’s ThistleCirsium tweedyi/eatonii – sprawls out of the rocks attracting a variety of pollinators. It is definitely one of the “good” thistles and is a native found only at high elevations.

 

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Fringed Grass-of-ParnassiaParnassia fimbriata – is worthy of a close look at the frilled petals, lobed yellow nectaries, sculpted pistil.  The oval leaves are also elegant.  These plants like moisture.

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Netveined WillowSalix reticulata – Willows are common and confusing in the alpine zones (and elsewhere!) This very low-growing species has small rounded leaves with distinctive veins (reticulate) making it is pretty easy to ID, I think…

Truly on the rocks:

 

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Miraculously, several plants cling onto rock faces such as found along the Rock Springs Loop trail below Cody Bowl.

 

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Brewer’s Cliffbrake FernPellea breweri – was a new one for me.  Black wiry petioles (stipes) hold out leathery evergreen leaflets.

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The black sori are protected by the folded leaf edges. Sori produce the spores that are key to reproduction in ferns.

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The pale lavender flowers of Mountain PenstemonPenstemon montanus – are fading, but are still a treat to see.  The plants are slightly woody and have toothed leaves (unusual for our WY species). The anthers are woolly.

 

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Common AlumrootHeurchera parvifolia – is a true rock lover! It is in the Saxifrage Family.  Saxi = rock

 

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Telsonix/Boykenia – Telesonix heucheraformis thrives in a rock crevice of a large glacial erratic on the way down to Cody Bowl. You may be able to see some family resemblance with Alumroot.

 

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Tufted RockmatPetrophyton caespitosum – forms a dense carpet draped over a ledge.

 

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No matter how you look at these plants, one can appreciate their adaptability and tenacity, growing in this challenging terrain—rock, snow, abrasive winds, drought, intense UV light, not to mention a growing season of maybe a couple of months.  And consider the luck of a seed to land in the right spot in the first place and mature and reproduce throughout many years of such trials.

If you don’t get up to admire these floral athletes this year, they will be there next year to enjoy on your alpine adventures.

Frances Clark, Wilson, WY,  August 24, 2019

*Note: Check on the trail conditions through the JH Mountain Resort before taking this hike.  At thie time of writing, the summit-access road was closed for construction projects. High elevation and rocky trail can be difficult for some hikers. Saying that we, were able to reach safely a wonderful display of flowers from the trail!

Flowers Blooming Low and High in Jackson Hole – a Sampling

All, it has been a busy wildflower season!  So many places to go and flowers to see.  Here are some observations of flowers around Jackson Hole during these last couple of weeks.  While fading at lower elevations, many of the same species are blooming, often with different companions, at higher altitudes and in different habitats. Others are more specialized to their particular niche.

Sageflats: Dry well drained soils and lots of sun!

TNP_VwTetons_frTSS_flws_7.10.19_1a_Q2_5x3_180A drive along Antelope Flats Road or inner Park Road rewards one with clouds of Sulphur Wild BuckwheatEriogonum umbellatumErioUmbe_fls_AntFlts_7.10.19_2_Q1_crp_5x5_180 Flowers go from white to cream to pink as they age.  The tiny, dried, winged fruits will be relished by rodents and birds.

Silky LupineLupinus sericeus – has been blooming for a week or two. Note the many hairs that cover the leaves and even back of the flowers.  The hairs help prevent water loss.  Bacteria are thriving in root nodules, providing the plants with usable nitrogen while the lofty lupine gives shelter and carbohydrates to the simple, tiny, yet crucial organisms.B_LupiSeri_habfl_AntFl_7513_2smIn some areas such as near Oxbow Bend or up Old Pass Road, Scarlet Gilia  — Ipomopsis aggregata — appears particularly abundant this year. In other areas, it stands more or less among sagebrush. IpomAggr_habfl_mass_OldPsRd_7.2.19_1_5x3_180Look for hummingbirds or sphinx moths visiting the red tubular flowers.  The long tubes are specialized to pollinators that can hover while drawing up the nectar deep inside the flower.  Flowers have no fragrance—most birds don’t smell but they do see red. IpomAggr_FlStCu_OldPsRd_7.2.19_1_Q2_crp_5x5_180These tubular “regular” (petals are all the same shape) red flowers in the phlox family are not to be confused with a more complex “irregular” flowers of the Wyoming Paintbrush — Castilleja liniariifolia, our state flower. Formerly placed in the Snapdragon Family, now Paintbrushes are in the Broomrape Family because they are hemiparasites on other plants, such as sage and lupines.  These plants derive sustenance, even chemical defenses, by attaching their weird roots to the roots of their hosts. CastLini_flst_Mung_6.24.17_1_Q2_crp_3x5_200The green Wyoming Paintbrush flowers lean out beyond the colorful red sepals.  Bracts just below add more color.  The leaves are also lean with narrow lobes. 

Hawksbeards – mostly TapertipCrepis acuminata – are abundant.  Look for the many (up to 100!) yellow flower heads, each with about 5-10 ray flowers, and 5-8 smooth “involucral bracts” below.  CrepAcum_flcuSide_20LkRd_71113_1_25x25Leaves have several deep, sharply toothed lobes and are a bit fuzzy or “tomentose”. CrepAcum_habfl_AntFl_62415_1bModoc HawksbeardC. modocensis – has fewer flower heads (up to 40/plant) but each has more than 10-60 ray flowers and the bracts are stippled with  black stiff hairs.  Leaves and stem are very hairy. See if you can find these two species and discern the differences.  There are other look-alike species as well.2b.CrepModo_fl_AntFl_52814_1crp180Q2_3x3

Another yellow composite with variations – Groundsels – rise about 12-18″ tall.  The bracts are all one size, smooth, often black-tipped, and contain yellow “ray” and “disk” flowers. This one is Rocky Mountain Groundsel –Packera. streptanthifolia. The leaves of three look-alike species are used for ID—leaves are variable in size and shape as they alternate up the stem.PackStre_habfl_TNP_PkRd_1_Q2_crp_3x5_180

Lance-leaved Stonecrops or SedumSedum lanceolatum – prefers well-drained soils to rocks. The pudgy leaves, which store water, alternate up the sprawling 4-6” stems.  SeduLanc_lfst_BTTr_62215_1-crp3x3

Flowers are bright yellow. SeduLanc_FlCU_SlgCrk_7.5.19_1_Q2_crp_5x3_180Members of the Stonecrop Family have a different type of metabolism – CAM – in order to do well in hot dry locations. They also are a host plant for the Rocky Mountain Parnassian butterfly Parnassius smintheus. (see: https://en.wikipedia.org/wiki/Parnassius_smintheus

Yellow Indian PaintbrushCastilleja flava – was still blooming strong in the dry sage and grasslands along Gros Ventre Road near the Red and Gray Hills. CastFlav_flhab_GVRd_7.11.19_1a_Q2_5x3_180Note the arrangement of its flowers:CastFlav_infl_bractCU_GVRd_7.10.19_1_Q1_crp_5x5_180

Also abundant in spots were bright magenta Northern Sweetvetch — Hedysarum boreale — clearly a member of the pea family. HedyBore_fls_GVRd_7.10.19_1_Q2_crp_5x5_180

Ballhead SandwortEremogone congesta – has a cluster of white flowers on the top of very thin stems.  Leaves are opposite and needle like. This is a common, if overlooked, wild relative to our store-bought pink carnations.EregCong_flst_AntFlts_7.10.19_1a_Q1_3x5_180

Various – and I mean various – fleabane daisiesErigeron spp.—are common.  They are hard to untangle botanically to this eye, in part because the roots are key to ID (and I don’t like to pull them up!), and then one examines hairs.  Just knowing it is a native daisy is good enough for me. And I don’t think the insects bother with the difference either.ErigPumi_flhab_MillB_6.1.16_1crp_5x3

Mountain Meadows – cooler and moisterMeadows around String Lake and up the trail to Ski Lake feature lush arrangements of taller “forbs” or perennial flowers:BTNF_SkiLkTr_vwDougFir_713.19_fix 5x3_180Sticky GeraniumsGeranium viscosissimum  — are abundant.  Here is a swallow-tail butterfly sucking up nectar. Anyone know which species?GeraVisc_fl_Swallowtail_GdwLk_7.14.19_1_Q1_crp_5x5_180Scarlet PaintbrushCastilleja miniata – is emerging. A_CastMini_fl_GoodLktr_7613_1Q2sm Note these flowers don’t lean out the same way the Wyoming Paintbrush does (compare with above photos) and the leaves are wider. Colors of paintbrushes can be variable but the flower details are relatively consistent…if they don’t hybridize with a nearby population of a similar species. Paintbrushes can be very confusing to ID!

Silvery LupineLupinus argenteus – is the common lupine of moist meadows and dappled light of aspen groves and evergreen forests.  The back of the “banner” petal is hairless, unlike Silky lupine which is silky-hairy on the back.  Hairs help reduce water loss, which is more of a concern in open drier sites. LupiArge_habfl_LSR_6.26.19_2a_crp_3x5_180

Tall CinquefoilPotentilla arguta – hold their flowers in tight erect fists.  The yellow-to-white flowers are held up on glandular stalks 2 or more feet high. PoteArgu_hab_191GrosVrd_71111_1a_3x5

These are often confused with the more broadly spreading flower clusters of Gland or Sticky CinquefoilP. glandulosa . Both are common (and sticky with glandular hairs). PoteGlan_flCU_PassTr_62915_1crpsmThese open, accessible flowers are important for a variety of pollinators.  Scientists have found plants contain a chemical that prevents tooth decay.   

Several blue penstemons –  Penstemon spp. — stand up through the vegetation or loom over a trail.  ID distinctions include the color, hairiness, length, and angle of the anthersPensCyacf_flside_SkiLkTr_7212_1Q2smBotanists and gardeners thrive on these minute differences.  Flower size affects bee pollinators which land on lower lip—they may or may not fit properly to reach the reward of nectar deep inside the flower while effectively carrying pollen to the next flower.PensCyan_flhab_SkiLkTr_7.12.19_1_Q2_crp_3x5_180This species found along the Ski Lake Trail keyed out to Wasatch Beardstongue — P. cyananthus.

Flowers of our perennial StickseedHackelia micrantha – mirror the summer blue sky above.  Later tiny fruits will stick to socks and fur. HackMicr_fl_SkiLktr_62815_1a_crp3x3

While phlox has mostly faded, its relative Nuttall’s Gilia — Leptosiphon nuttalli – (no good common name) is beginning to bloom.  The deeply dissected leaves form frilly whorls up the slightly woody stems that form a 12″ tall mound. 29.LeptNutt_lfflCU_SkiLktr_62815_1a_Q2_crpCU_1.5sq_300Flowers are extremely fragrant.  Bend over for a whiff.  It is also related to Scarlet Gilia.

Western SweetrootOsmorhiza occidentalis – is common in meadows to open forest. OsmoOcci_Flst_HistTr_7.2.19_1_5x3_180The tiny yellow-green flowers arranged in umbels will hold up schizocarps (split fruits) that taste like licorice.  It is related to cumin, coriander, dill, anise, and many other tangy herbs in the Parsley family. 

Louseworts are going and coming.   Fern-leaf LousewortPedicularis bracteosa – is fading fast at lower elevations, but beginning to bloom over 8,500’. PediBrac_flhab_TetPsS_7.3.18_1b_Q2_5x3_180smIn lodgepole pine forests, Parrot’s BeaksPedicularis racemosa – are unfurling their flowers to attract specific pollinators.  Usually a smart bee, channeled by nectar guides and the unique shape of the flower, lands on the lower petals, wriggles around, and vibrates its wings.  Pollen grains bounce out of the beak-like tube of the upper petal and attach to the bee’s hairy back.  Even with its long-comb-like legs, the bee can’t glean all the pollen: some remains out of reach in the crevice between the bee’s head and thorax.  Visiting the next ready flower, the bee’s positioning will cause the stigma to curve around and tap the pollen out of the crack for successful fertilization! (see lower right flower)PediRace_fl_SkiLktr_62815_7crpsm

High Elevations – late blooming!

Elevations above about 8,500’ are noticeably delayed in flower this year.  South of Teton Pass and above Ski and Goodwin Lakes, one finds only early spring flowers.  Carpets of Spring BeautiesClaytonia lanceolata — are sprouting where snow drifts are finally melting. ClayLati_habfl_mass_GdwLk_7.14.19_1a_Q1_fix_crp_5x3_180Species particularly adapted to very short growing seasons are also blooming where snow has just melted.  Patches of Rocky Mountain Snow ButtercupsRanunculus adoneus – are frequent. Note their fine leaves.RanuAdon_hab_mass_SkiLkTr_7.12.19_2a_Q2_5x3_180.jpg

There are many, many more flowers to see.  This is just a preview and hopefully incentive for you to explore, identify, and understand the remarkable plant world around us.

Frances Clark, Wilson, WY

July 16, 2019

We appreciate comments, corrections, and additions. Please let us know what you think about this posting.  We will do our best to incorporate your thougts.

 

Tra-la – It’s May! – Early Spring Flowers 2019

WyHab_WL_Mosaic_GameCrk_42915_5x3_180.jpgSpring sun is warming south-facing slopes of buttes and hillsides.  Snow along Grand Teton National Park roads is finally retreating. Wetlands are warming. Bugs and birds are flying about. The delight is in the details of small flowers; no big show yet.

Along roads and low openings in the valley:  

Patches of yellow buttercups are the first to draw the eye.

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ButtercupRanunculus spp. – flowers gleam, and uniquely so. The outer layer of the petals – epidermis – is only one-cell thick and the cells are particularly thin and flat.  They hold pigments that absorb blue-green wave lengths of light. Thus yellow wavelengths can keep going through the outer cell layer, penetrate a thin air layer, and then reach a starch layer of cells that scatters the yellow light back up through the pigments again. Furthermore, the thin outer layer with air layer just beneath has the physical properties of a thin-film, creating the shiny look to the flowers. The combination os pigmentation and unique structural qualities of the flower cells provide the bright glossy yellow found only in buttercups and a few cousins.  At certain angles, flowers actually flash a signal to passing pollinators to come visit. (For much more to this complex story see references below.)

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Buttercup petals beam intense yellow light and UV wave-lenghs that signal to pollinators. Also, the curving petals with the special cell layers provide addtional warming to the reproductive parts in the center, speeding up the life cycle.

Look closely at our two similar species of buttercups:

RanGlab_habfl_AntFlts_5411_5Q1smThe lowest leaf of Sagebrush ButtercupRanunculus glaberrimus – is unlobed, the upper leaves are 3-lobed. It is a denizen of sage flats.

RanuGlabcf_flhab3_nopetals_BTBut_5.6.19_1_Q2_crp_3x5_180Some individual Sagebrush Buttercups don’t have petals, only sepals. I dont’ know why the flashy petals aren’t there.

RanuJovi_flhab_Cluster2_MuriRch_4.21.18_1aQ2_4x3_180fix.jpgIn Utah ButtercupR. jovis – both the lower and upper leaves are lobed into three parts. Note buttercups have many separate anthers and stigmas—a common characteristic of this family. It is found in relatively moist locations, including woodland edges and openings.

ClayLanc_fllv_JosRdg_2614_1crp180sm.jpgSpringbeautiesClaytonia lanceolata — grow in scattered in patches. Some blooms exhibit obvious pinkish veins that direct pollinators to yellow nectaries in the center. Pollinators bump against the anthers and get dusted with pollen.

OrogLine_habfl_RkCrk_5.2.19It is easy to step on Turkey PeasOrogenia lineariifolia. The plants look like bits of lichen or stone, nothing to think about.

OrogLini_fl_Elkscat_MuriRch_4.21.18_3_crp_5x3.5_180However, Turkey Peas are more interesting if you take a close look at their tiny white flower with maroon centers that together form clusters barely an inch long. Think about what tiny insects must pollinate them–likey small flies and bees.

OrogLin_bulbCU_WilBrk_4912_1a_5x3-180.jpgSandhill cranes, bears, and rodents seek out the thumb-sized bulbs (“peas”) for food.  (Turkeys would likely eat the bulbs if they lived in Jackson.)

DiceUnif_habfl_PkRd_4.23.19_crp_5x3_180ipThe quintessential western plant Steer’s-headDicentra uniflora – requires some belly botany. Scan an area for divided leaves and then get down to stare at the steer-like flowers. This is the larval host plant for the Parnassian butterfly Parnassius clodius, which Dr. Debrinski from MSU is researching in Grant Teton National Park (more info on her research below).

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Parnassian butterfly species on Blacktail Butte 5.4.19.

 

FriPudi_fl_SchwLd_bench_fl_52011_1a_5x3_180fixYellowbellsFritillaria pudica – are always cheerful! The 6-8”-high plants sprout from miniature scaly bulbs. The base of the 6 yellow tepals is said to change from red to green depending on pollination, but I can’t see any consistent difference happening to the outside flower color or anthers and pistil on the inside. Maybe you can.

Violets are flowering here and there:

ViolPurp_habfl_RkCrk_5.2.19_1_crp_5x5_180Goosefoot VioletViola purpurea var. venosa – has leaves shaped like goose feet with a few more toes. The back of the leaves and yellow petals are often purple, hence “purpurea” in its botanical name. Note the dark center of the flower and the convenient landing pad of petals for pollinators.

ViolNutt_fl_JosieRidge_fl52011_1crp180sm.jpgSeveral yellow violets intergrade in leaf features which confuse me and other botanists trying to sort out the names.  This cheerful specimen is one of three look-alike species – V. vallicola, V. praemorsa, or V. nuttalii.  Leaf ratios, shapes, and hairiness, as well as ultimately seed-capsule sizes, determine identification.

Dry slopes and knolls:  Rambles up the south side of Blacktail Butte and rocky knolls around Kelly Warm Springs yield treasures tucked into the rocky soils. Many of the plants are silvery and/or hairy and grow very slowly into low mounds or mats—adaptations to limited water and nutrients and intense light and wind.

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Please watch you step…these small plants can be hard to see and some are very old.

PhloHood_flhab_BltB_4.13.17_1_5x3-180sm.jpgHood’s PhloxPhlox hoodii – is often the first out, with its white to bluish flowers. Bees and flies pick up on the sweet fragrance. They come in and land on the flared petal tips and dip their long tongues deep down the center tube for nectar. They then carry the orange pollen off to other flowers nearby. The leaves of Hood’s Phlox are opposite, very small and tight on very slow growing stems that collectively form a cushion shape.  Plants inches wide can be decades old.

WyPl_PhysDidy_FlhabCU_Bt_3.22.15_1Q2_5x3_180Nearby, TwinpodsPhysaria didymocarpa – feature bright-yellow, 4-petalled flowers at the end of sprawling 3-4” stems. Spade-shaped, silvery leaves help identify this member of the Mustard Family. Mustards usually have 4 petals, 6 anthers (2 short, 4 long), and one 2-parted pistil.

AnteDimo_flfm_lvs_KWmSp_5.4.19_1_Q2_crp_5x3_180The first pussytoes to bloom is Low PussytoesAntennaria dimorpha.  The tiny gray, finely hairy leaves form mats on the ground. Look closely for the flowers.

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Individual female flowers have stigmas…

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that fork to capture pollen.  No males in sight.

Note female and male flowers are on separate plants. This separation helps encourage cross-pollination, but if there are no pollinators present, females can set seeds on their own. Female plants often outnumber male plants in a population.

A_CymoLonp_habfl_kws_42614_3Q1_crpsm180.jpgSprawling CymopterisCymopteris longipes – is spreading its whorl of dissected silvery leaves low to the ground.  As a member of the Carrot Family, plants have umbels, in this case with yellow flowers.

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Cymopteris longipes has a buried pseudoscape. This buried stem is surrounded by old leaf bases. The true roots actually branch from below the area shown here.

Later, the underground stem will stretch up lifting the leaves higher to the light above growing competition. The stalk of the umbel will extend, too, elevating the winged fruits into the wind mid-summer.

AstrPurs_flhab_BTBut_5.6.19_1_crp_5x3_180Pursh’s MilkvetchAstragalus purshii – is also just beginning to flower on dry knolls.  The pea-like flowers are slightly yellow to white with a blue bow to the keel (lower two petals). Some flowers open wide for pollinator business. Note the pinnately divided leaves are silvery hairy.

CastChro_bractsLvs_BTBut_5.6.19_1_5x3_180The brilliant red of paintbrushCastilleja chromosa – is provided by the leaf-like  bracts. Soon tubular flowers will emerge from their axils. I am not sure why there is so much color without the presence of any flowers yet. Maybe the plants are announcing to pollinators: opening for business soon!

TownLepi_fllv_KWmSp_5.4.19_1a_Q1_crp_3x3_180Our local Townsendias belong to a beautiful but often confusing genus. This plant has all the features of T. leptotes: narrow leaves, whitish petals, a whorl of 4-5 rows of pointed bracts tinged with color. Apparently this species and T. montana can hybridize or self-fertilize to the point that some experts say separating the two species appears “arbitrary.” I say, let’s just enjoy the flowers if you can find them. They are pretty rare.

Wetlands with catkins:

AlnuInca_flm_MWRd_4.22.19_1a_Q2_5x3_200.jpgShrub swamps throughout the valley are warming up. Ducks, moose, and beaver are moving through the waters under dangling catkins of alders and amidst thickets of pussywillows.

AlnuInca_flMfm_FallCrkRd_4.29.18_1_Q1_3x5_180Male catkins of mountain aldersAlnus incana var. occidentalis – elongate: their pollen is released upon the wind to meet up – purely by chance – with the stigmas of female flowers (above left in photo) in separate, stout “cones.”

BetuOcci_Flst_CUGameCrk_5.30.18_1_Q1_crp_5x3_180Later in May, its relative Bog BirchBetula glandulosa – will bloom after its leaves have filled out.

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Male catkin of Booth’s Willow – Salix boothii.  Notice the waxy “bloom” on the greenish yellow stem that can rub off.  This is a helpful winter ID feature.

Willows (Salix spp.) of various kinds (and there are many) are bursting their buds and producing male or female “pussies”.  In willows, female fruits (capsules) are the definitive for identification, but are often elusive. Bees pollinate many willows—they seek out nectar at the base of tiny, petal-less flowers.

WyPl_PopuSp_BudMCU_AntFl_42415_6bQ2_crp_5x3_180

A male catkin of cottonwood – note the red anthers that will soon shed pollen.

Cottonwood and its congener aspen (both are in the genus Populus) also have catkins, again males and females on separate plants.  It is fascinating to investigate the differences.

We are seeing just the first flurry of flowers.  We will try to keep you posted on new arrivals.

Enjoy your adventures into spring!

Frances Clark, Teton Plants 5.7.19

P.S. We always appreciate comments and corrections.  Please send an email to tetonplants@gmail.com

References:

“How Buttercups Get Their Gloss” by Stephanie Pappas, Live Science Contributor – link: https://www.livescience.com/57964-how-buttercups-get-their-yellow-gloss.html

“Scientists Discover why buttercups reflect yellow on chins”. By University of Cambridge, Phys.org.  December 2011. https://phys.org/news/2011-12-scientists-buttercups-yellow-chins.html

“Functional optics of glossy buttercup flowers” by CJ van der Kooi, et al.  Journal of Royal Society Interface 14.  Fascinating details including photos of the physics. Available on line at  https://royalsocietypublishing.org/doi/full/10.1098/rsif.2016.0933

Dr. Diane Debinski is studying Clodius Parnassia butterfly populations in Grand Teton National Park. Here are a few links to her research:

https://lib.dr.iastate.edu/nrem_pubs/274/

https://repository.uwyo.edu/uwnpsrc_reports/vol38/iss1/12/

Time to Hike for Subalpine to Alpine Flowers before they Fade

BTNF_Rendv_CodyB_VwNUp_8.7.18_1s_Q1_5x3_180Wildflower seekers are hiking above 8,000’, even 9,000′, for colorful displays of flowers found earlier at lower elevations.   It is also time to search for unusual subalpine to alpine flowers above 9,500′ to 10,500′

Here are some recent sightings and identification tips, focusing on species specialized to grow in the very short summer season, instense sun, abrasive wind, and poor soils of high altitudes. Most of the photos were taken in the past week up on Rendezvous Mountain, but the species can be seen elsewhere.

The majority of the photos below were taken in the last 10 days around Rendezvous Mountain. The photos focus on identification tips. You are welcome to ignore the details and just enjoy their beauty. If you are a full flower nerd, know that the taxonomists do not always agree with the classification of some of the species shown…opinions among scientists often vary.

Starting from the top:

HappSuff_SympFoliApr_habfl_Rendv_8.11.18_1_Q2_fix_5x3_180

Woolly GoldenweedHapplopappus suffruticosus – is cheerfully sprawling along the road down Rendezvous. These subshrubs have woody bases, alternate twisted leaves, and bright yellow flowers (photo above and below).

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Eaton’s/Tweedy’s ThistleCirsium eatonii/tweedyi. – is a 2-4’ tall, native (good!) thistle which catches your attention:

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Carefully, look into the dense cluster of flowers at the top of the plants: The bracts are interlaced by a web of glistening hairs. Bracts surround 2-3” pinkish flower heads. Pollinators are plentiful!

CirsTwee_flBracts_Rendv_8.6.18_1_Q2_5x5_180Leaves are 6-8” long, toothed to lobed, wavy, and spine-tipped. Notably, petioles run down the stem (decurrent). Tweedy’s thistle is deemed “unresolved” by the authoritative Flora of North America: it is not even considered a variety. However, the authors note that there is much post-glacial hybridization among formerly isolated populations of this complex genus.

ArniLong_habfl_Rendv_8.11.18_1_Q1_5x3_180Bright yellow Long-leaved ArnicaArnica longifolia – grows in cheerful drifts.

ArniLong_flCU_Rendv_8.6.18_1_Q1_5x3_180The 1-1.5’ stems feature several pairs of elongate leaves, as well as many yellow heads surrounded by equal-length bracts. The plant is overall glandular hairy with a strong odor.

ArniLong_habfl_Rendv_road_8.11.18_1_Q1_5x3_180It is common as you continue down the mountain road and seen from the tram.

As you head down farther where snow piled high in the lee of mountain slope or in a bowl, plants are larger and more profuse. Here a few showy species seen on the way down to Cody Bowl:

BTNF_Rendv_CodyB_VwNUp_8.7.18_1s_Q1_5x3_180

HedyAmerAlp_habfl_Rendv_8.11.18_1_Q2_fix_5x3_180Alpine sweetvetchHedysarum alpinum var. americanum – has many deep violet- red, pea-like flowers (its in the pea family) dangling from one side of the inflorescence (photos above and  below).

HedyAmerAlp_fl_Rendv_8.11.18_3a_Q2_5x5_180Bluish-green leaves are pinnately divided into oval leaflets. Careful measurements of flowers (9-15mm) distinguish it from the slightly larger flowered (17-22 mm) Western Sweetvetch (H. occidentale). (Not all taxonomists agree with this separation!)

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The fruits of Sweetvetches are termed loments – segmented fruits each with one seed inside. I think of them as pods flatted by a steam roller. The flat segments break off and disperse upon the wind. Many are forming now (photo above).

White-coiled LousewortPedicularis contorta – has distinctive “coiled” or beaked white flowers which have evolved to fit worker bumblebee pollinators (below). Note the stigma projecting from the coiled  beak formed by fused petals.  When a bee lands, the stigma fits between the bee’s head and body and picks up pollen which the bee could not reach from a visit to another lousewort flower.

PediCont_flhand_SkiLktr_62815_4.a_Q2_5X5_180The 1’ plants have divided, mostly basal leaves, which helps distinguish it from a similar, more lower-elevation species: Parrot’s Beak – P. racemosa – which has with toothed but not lobed leaves.  White-coiled lousewort is shown  below.

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Sulphur PaintbrushCastilleja sulphurea – continues to bloom in the shade of Engelmann spruce and Subalpine Fir, where snow collected over winter and lasted longer.

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GroundselsSenecio/Packera  – are common and confounding yellow composites. They have equal length bracts like Arnicas, but the leaves alternate up the stem.

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The relatively large Thickleaf GroundselSenecio crassulus – is still blooming strong in mountain meadows, such as Rocky Springs Loop (above). Look for the even-sized, waxy bracts with black somewhat frayed tips.

SeneCras_flCU_PassTr_71615_1_Q15x5_180_Undulating, slightly toothed, slightly succulent or waxy leaves clasp the 1-2’ stems (below).

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Related and once grouped in with Senecios, two different “Packeras” continue to challenge this and other botanists. The expert Arthur Cronquist said groundsels are a “transcontinental complex of ill-defined taxa.” It is a challenging (frustrating!) botanical puzzle to try to tell them apart!

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Rocky Mountain GroundselPackera/Senecio streptanthifolia – is found frequently at lower elevations, and more occasionally at high elevations. The lower leaves are toothed or lobed more or less, as are the upper leaves. The lower leaves are larger than the upper leaves. Plants have many yellow flower heads. I am not sure which species this is! (photo above).

Different GroundselPackera/Senecio dimorphophylla – is often hard to tell apart from Rocky mountain groundsel. However the scientific name di- (two), morpho- (shape), phylla (leaf) indicates a distinct difference in the shape and size of basal vs. stem leaves. The basal leaves tend to be unlobed to toothed, the upper leaves more deeply lobed and clasping the stem with arrow-shaped leaf bases (auriculate). At least a few stem-leaves are often larger or equal in size to the basal leaves (I have noticed that the lowest stem leaf is often the largest and most indicative of the species).  The photo below seems a clear identification of this species. I also noted that it seeds in readily to the harsh conditions of talus. PackDimo_flhab_Rendv_8.6.18_1_Q1_3x5_180

Asters are beginning to appear.  A common species at high elevations is the Leafybract AsterSymphiotrichum foliaceum var. apricum – which is low growing, with slightly clasping leaves, and wide bracts (foliaceous) with purple tips surrounding the lavender heads (photo below).

SympFoliApri_FlCU_RkSprLp_8.11.18_1b_Q1_crp_3x5_180

Thickstem asterEurybia integrifolius – also has flaring, variable bracts. Flower heads are sticky hairy and a distinctive deep violet blue. Reddish stems are also glandular hairy. This 1-2.5’ plant grows readily at lower elevations, as well (photo below).EurInt_fl_MwRd_9611_q2crpsm

Always a special treat to see, Mountain Bog GentianGentian calycosa – appears particularly abundant in moist to wet sites down to Cody Bowl. Observe the delicate lines and spots that attract bumblebee pollinators deep into the open bell shaped flowers. In this photo (below), gentian is nestled in with the common Rocky Mountain Goldenrod.

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Rocky Mountain GoldenrodSolidago multiradiata – is a common plant at both high and lower elevations in Teton County.  To tell it apart from other golendrods, look for clusters of leaves at the base and stiff hairs along the petioles of the lower leaves –a sure way to know the species.  The heads also have over 13 tiny ray flowers – the “multi-radiata” in its name.  Height varies from an inch to a foot or so, depending on the conditions it is growing in.

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BTNF_Rendv_CodyB_vwtalus_up_8.6.18_1s_Q2_5x3_180The rocky talus of Cody Bowl has several speciality flowers that seem to be able to grow out of rock.SeneFrem_flhab_pol_Rendv_8.6.18_1a_Q2_5x3_180Fremont GroundselSenecio fremontii – has single flowers with the indicative row of waxy bracts. The small leaves are toothed and arranged along the stem: not clustered at the base (although some plants with short stems look like they have basal leaves).  The flowers are single and appear large compared to the leafy body of the plant.

One unusual species Alpine GroundselLigularia/Senecio amplectans – is also part of the groundsel group: Most notable are mostly solitary (1-3), nodding, ½” heads. The leaves are mostly basal and fine-toothed.

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Other species are much easier to identify and appreciate:

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Parry’s PrimrosePrimula parryii – grows very rarely in the talus of Cody Bowl. I have also seen it near the top of Targhee in a similar rocky habitat.

AnemParvi_flhab_Rendv_8.6.18_1_Q2_5x5_180Small-flowered AnemoneAnemone parviflora – is only inches tall with tidy whorled leaves.

AnemTeto_flhab_Rendv_8.6.18_1_Q2_crp_5x5_180And nearby, the deep-pink Teton AnemoneAnemone tetonensis.

AnemSp_frCU_Rendv_72415_1_5x5_180Some anemones are already setting seeds!

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A particularly tiny plant: Alp LilyLloydia serotina – is only 2-6” high. Its 6 white “tepals” remind us it is in the Lily Family. It grows from bulbs.

CampUnif_flhab_pol_Rendv_8.6.18_1_Q2_fix_5x5_180Similar in size is the Alpine HarebellCampanula uniflora. Instead of many bell-like flowers per stem found in the more common harebell, this species has only one flower per stem, as the Latin name uni-flora aptly describes.

PoleVisc_flhab_hand_Rendv_8.6.18_1_Q1_5x3_180You may smell this plant before you see it’s blue flowers: Skypilot or SkunkflowerPolemonium viscosum. It ranges in size from 5-12” high.

As one heads down into the lower portion of Rock Springs Loop, flowers once again become plentiful:

BTNF_Rendv_RkSprLp_vwFlmix_8.11.18_Q2_fix_5x3_180Snow ButtercupsRanunculus adoneus – are blooming brightly in recently melted snowpatches. The leaves are divided 1-2x into narrow lobes (photo below).RanuAdoe_flhab_Rendv_8.6.18_1a_Q1_5x3_180

A flower mix of more subalpine or meadow species is patchy on the steep slopes.  Below is a combination of Nuttall’s LeptosiphonLeptosiphon nuttallii – and very low growing Thickleaf GrounsdselBTNF_Rendv_vwRockSpLp_SeneCras_LeptNutt_8.11.18_1_Q1_fix_5x3_180Another combination includes Sulphur paintbrush, Bog Gentian, and Coiled-beak Lousewort:BTNF_Rendv_vwRockSpLp_flmix_GEntCaly_LeptNutt_8.11.18_2a_Q1

There is much more to discover in the high elevations before snow flies. Enjoy looking for the above species and more these last few weeks of summer.

And for other flowers still blooming at this time and with more aster ID go to our 2016 archives: “Get High on Wildflowers”:  https://tetonplants.org/2016/08/

Frances Clark, Wilson, WY

Corrections and comments welcome!

 

 

 

What Blooms in Wildfire Burns?

RkPkwy_Burn_PeriMont_vw4_8.2.18_1_s_Q1_5x3-180Forest fires can appear devastating at first, but for the most part nature has its systems for resilience. Depending on how hot the fire was and what plants were present both above and below ground and nearby, vegetation will return in its own due course. In some cases, plants sprout that have not been noticed in years, and indeed are triggered to flower after the heat of the moment. Others take advantage of the open ground and fly in with fresh seeds. Still others have stored seed until the magic moment. Wildlife also takes advantage of the changes.

Lodgepole PinePinus contorta – is a fire-adapted species. While the thin-barked trees are killed, thick “serotinous” cones have held seeds for years.PinuCont_FrSerCU_MWRd_32212_1.jpgTheir cones have thick scales with spine tips which protect the seeds inside from mauraders and weather for years.  When a fire comes through, the resin that has sealed the scales shut melts, and cone scales open wide, releasing winged seeds upon the wind. The delicate embryos fall onto newly exposed soil, which may be enhanced by ash, and quickly germinate. Ash often contains recycled nutrients and retains warmth which helps the seeds grow.  Seeds germinate quickly, giving them a headstart among competing plants.  Pines in fact need sun to grow well. A truly fire adapated species!

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Mountain MallowIlliamna rivularis – often appears in great numbers after a burn.  Affected by extreme heat, their thick seed coats crack, enabling seeds to imbibe water and sprout. These seeds may have lain buried for decades in the soil waiting for such a moment. BTNF_PalmCanTr_Burn_IlliRivu_7_7.13.18_Q2_5x3_180Due to a prescribed burn south of Hoback, the slopes along the trail up Palmer Creek are now covered with 4-5’ flowering Mountain Mallow plants (photo above taken 7.13.18). Soon fruits, which look like peeled hairy tangerines, will split to release seeds for the next generation decades in the future (photo below).IlliRivu_frsSt_LeiLk_91213_2aCrpsmNote: In mountain mallow the seedbank is in the soil, in lodgepole pine, the seedbank is in the air.

Another plant that responds uniquely to fire is SnowbrushCeonothus velutinus. Hikers can see a profusion of Snowbrush along String Lake (below) and on the way to Taggart Lake in Teton National Park.TNP17_StrLkTr_CeonVelu_CU_WyHab_6.30.17_2_5x3_180This evergreen, resinous, sprawling shrub will shoot up new branches from old roots after a light fire. After heavy burns, it can also sprout from “Rip-van-Winkle” seeds.CeonVelu_fllfCU__StrLk_71105_2_3x1_180

Flowers blooming almost a century ago produced seeds that have been lying in wait until heat and sun stimulated them to germinate. CeonVelu_frLvs_BTTr_82013_1_5x3_180

Others report a profusion of White SpiraeaSpiraea betulifolia – blooming (photo below) within the 34,000-acre area of the Cliff Creek Fire, also of 2016.  This appears to be another species is “released” after a fire.SpirBetu_fllf_20LkRd_71113_1a_5x3_180.jpg

The results of the 20,000+ acre Berry Fire are visible from the Rockefeller Memorial Parkway (photo below) and Grassy Lake Road. The 2016 fire burned fast and hot in some areas forming a mosaic of impact.RkPkwy_Burn_PeriMont_8.2.18_5a_s_Q2_5x3_180Notably, in some areas it burned through lodgepole stands that were recolonizing from a fire only a few years before. Ecologists and foresters are concerned that this unusual short “return” interval will be the pattern of future fires in this era of climate disruption.

PinegrassCalamagrostis rubescens – is a tufted, long-leaved grass that rarely blooms. While a common groundcover in the shade of a forest, it usually goes unnoticed by hikers because it is “just a grass.” However, with the stimulus of fire and sun, 2-3’ stalks of delicate flowers shoot up and flourish (photo below). TNP17_Burn_CalaRube_8.17.17_2_Q2_5x3_180 Deep fibrous roots of Pinegrass are important for holding soils, especially when soils are vulnerable to erosion after fires. Plants are blooming in profusion near the parkway.

RkPkwy_Burn_For_flwMix_Epil_Cala_8.2.18_1_s_Q2_5x3_250FireweedEpilobium/Chamerion angustifolia – is well known for showing up after fires. In the insulating soil, rhizomatous (underground creeping) stems growing 4-6” deep may have survived the above-ground heat to sprout again. Even one surviving plant can shed 1000s of seeds that can catch upon the wind, land, and germinate quickly on exposed ground. (Photo above shows both Fireweed and Pinegrass.)

Other plants flowering among blackend trunks include several members of the Aster Family which have deep roots and seeds dispersed by wind.RkPkwy_Burn_ArniXdive_8.2.18_2_s_Q2_crp_%x3_180Cheerful patches of Broadleaf ArnicaArnica latifolia – and a strange hybrid, likely Arnica X diversifolia – a cross between Heartleaf and Broadleaf arnica, are growing in charred soils (photo above).RkPkwy_Burn_GrsLkRd_SoliMiss_8.2.18_2_s_Q1_5x3_180Large clumps of yellow Missouri GoldenrodSolidago missouriensis – was dense along Grassy Lake Road, brightening the dark scene (photo above).RkPkwy_Burn_GrsLkRd_flwMix_EuryInte_Achi_8.2.18_2_s_Q2_5x3_180A mix of YarrowAchillea millefolium – and Thickstem AsterEurybia integrifolia are common in fields right now, but they are also flourishing in the sun under dead lodgepole pine trees along Grassy Lake Road (photo above).

TNP17_Burn_LupiArge_8.17.17_1_5x3_180Silvery LupineLupinus argenteus – seeds are “scarified” by the heat of fire, enabling  buried seeds to germinate relatively quickly.  As a legume, lupines have a mutually beneficial relationship with bacteria in their root nodules that can “fix” nitrogen. This provides lupines an advantage in colonizing poor soils (photo above). Their heavy seeds pop out of their pea-pod like fruits.RkPkwy_Burn_DracParvi_hab_8.2.18_1_s_Q2_crp_5x3_180A robust member of the Mint Family – DragonheadDracocephalum parviflorum – (photo above) was a new species to this botanist. Apparently it thrives in disturbed soils.RkPkwy_Burn_flwmix_ErigSpec_Peri_Lupi_8.2.18_1_s_Q2_5x3_250Patches of other common meadow flowers have retained a niche as well, including Oregon DaisyErigeron speciosus – (photo above) with its many narrow, lavender ray flowers (ray flowers look like petals). Many perennials have deep storage roots that are often insulated by soils to heat of fire (or the cold of winter.)

It is unclear to this writer how much of the open areas between forest patches of the Berry Fire actually burned, if at all.  (Do you know?) Often meadows don’t provide enough fuel to carry a fire. However, embers often fly across roads, wetlands, and meadow, igniting trees despite the intevening “fire breaks.” In any case, this is what is growing in the meadows.

RkPkwy_Burn_PeriMont_vw1_8.2.18_2_s_Q1_5x3_180Common YampaPerideridia montana – has created a tapestry of white. Upon a walk through the area, one can see that many late-summer flowers which are common elsewhere as here as well: a hidden layer of Sticky GeraniumGeranium viscossimum, blue spires of Tall DelphiniumDelphinium occidentale, yellow sprays of CinquefoilPotentilla spp., orange-yellow Rocky Mountain GolendrodSolidago multiradiata, and spikes of blue Silvery Lupine mix in.

Common grasses include: stiff spikes of TimothyPhleum pratensis (photo below), PhlePrat_fl_2OL_8714_3_5x5_180Mountain BromeBromus carinatus (photo below),BromCari_fl_BTTr_62715_1_5x5_180and elegant spikelets of OniongrassMelica spectabilis (photo below):MeliSpec_flCU_BTTrHd_62215_2Q2_5x5_160(Note all the grasses pictured above are in bloom)

These grasses have dense deep roots or bulbs, as in the aptly named Oniongrass (below).MeliSpec_bulbfl_SkiLktr_62815_1acrpsmGrasses have evolved to sprout from buds at the base of their leaves – an adaptation to both browsing and fire.

As for wildlife, signs of elk are frequent–they enjoy nutritious grasses. Bears will enjoy the storage roots of yampa come spring—or perhaps pocket gophers, which also eat yampa roots. A week ago, a pair of Sandhill Cranes was walking through the downed trunks, feeding on insects. Black-backed and American Three-toed Woodpeckers seek out burned-over forests as long as the bark remains. Using their chisel-like bills, these woodpeckers feast on insects feeding and breeding under peeling, split bark of weakened or dead trees.

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Despite the stark appearance, all is not lost after a wildfire.TNP17_Burn_Logs_EpilAngu_8.17.17_1_5x3_180

Much is being researched and understood about fire ecology.  It is facinating to conduct your own observations.  We have a wonderful opportunity to see the variations in progression at the Berry and Cliff Creek Fires, both of which were started by lightning two years ago.

Frances Clark, Wilson, WY

Meadow Flowers: Obvious Favorites of the Sun

BTNF_Munger_vwFl_7.4.18_3_q2_5x3_180smWith summer strong, flowers are blooming everywhere in and around Jackson Hole. Here are a favorite dozen (plus!) wildflowers seen on hikes this past week: around Phelps Lake and up Munger Mountain (above); the start of the Ski Lake Trail; just south of Teton Pass; and the north end of Grand Teton National Park.  Lower elevations fade first while upper elevations are just emerging.

As always, it is fascinating to observe flower shapes and color and to discover which pollinators come to visit. Much is still unknown about how flowers work. Also, much of the action occurs underground.  Enjoy your investigations.

Sticky Geraniums (Geranium viscosissimum) are abundant both in full sun and under the shade of aspen trees. Their wide-open pink to magenta flowers attract pollinators of various sizes. Nectar guides—dark lines—lead into the center of the flower to the reward of both nectar and pollen. Male pollen is offered first by 10 anthers, and as the flower matures, five female stigmas are then exposed to gather pollen from insect visitors.  This way it is not fertilized by itself, which can cause inbreeding depression.

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Wide open flowers invite a variety of pollinators.  Here the male anthers are ready before the femail stigmas which lie in the very center of the flower.

One-flowered Little Sunflowers (Helianthella uniflora) form masses of cheerful yellow on hillsides.

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About 2-2.5’ tall, each of its many stems sports more or less opposite leaves with three strong veins. The stems are topped off with at least one 2”-wide flower head.

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Here the Little Sunflowers are flourishing high on Munger Mountain July 4.

Its more robust relative Five-veined Little Sunflower (Helianthella quinquenervis) stands taller and glares right at you.

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Note the number of pollinators and the individual small flowers beginning to bloom!  As composites, sunflowers have many tiny flowers that unfurl in a spiral, starting on the outside.  These many flowers form a “head”.

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The large lower leaves have 5 distinct nerves (quinque = 5 and nervis = nerves) and plants usually have only one big 3-4” flower head per stem.

Both of these sunflowers are relatively small compared to cultivated sunflowers (Helianthus annuus) produced for oil and seed. Note, a sunflower head has many tiny flowers that bloom in a spiral sequence. Each flower will produce a fruit with a single seed—think about unshelled sunflower seeds—the shell or husk is the fruit, with a nutritious seed inside. Birds will flock to the seeds when ripe.

Fernleaf Lovage (Ligusticum filicinum) or osha is just coming into flower in some places. Individual tiny flowers are held out in umbels—structures similar to ribs of an umbrella. Umbels are a distinctive feature of the Parsley or former Umbelliferae Family.

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Lovage leaves are finely dissected, similarly to its relative—carrot or Queen’s Anne’s lace.

Osha

Below the frilly skirts of leaves grows a dense collection of pungent roots that Native Americans have used for centuries for medicinal purposes.

Giant Hyssop (Agastache urticifolia) is one of the few members of the fragrant mint family in Jackson Hole.

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Like all mints, the 2-4’ stems of Giant Hyssop are square, the scented leaves are opposite, and the flowers “bilabiate” or “irregular” e.g. flowers have two similar halves—like our faces.

Long anthers stick out, distributing pollen on the heads of hovering hummingbirds or on bodies of pushy bees which use the lower petals as platforms.  Upon visits to other hyssop flowers, these pollinators distribute pollen to female parts which form seeds.

While several of the flowers listed below have faded in southern, lower reaches of Jackson Hole, they are blooming abundantly up near Oxbow Bend and at higher elevations.

TNP17_vw_Tetons_Antflt_frTSSrd_6.19.17_1c_sm5x3_180

Many flowers that bloomed around Antelope Flats a few weeks ago are now blooming at higher elevations or more northern reaches of the park.

Sulphur Buckwheat (Eriogonum umbellatum) forms “clouds” of flowers above mats of ½” oval leaves.

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At this time of year, 1’ stems shoot up forming clouds of fluffy creamy white to yellow to rose flowers. They float low over the hillside or sageflats.  At the base of the “inflorescence” is a whorl or collar of oval leaves.

Sulphur Buckwheat flowers provide valuable nectar to pollinators, such as Parnassian Butterflies. Dr. Diane Debinski of Montana State University is investigating the relation of this species and Clodius Parnassian butterflies (Parnassius clodius) near Pacific Creek to determine impacts of climate change on insect populations.

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I think this is a Clodius Parnassian butterfly which is being studied.  The species nectars on Sulphur Buckwheat flowers.

Towering up between sulphur flowers, wands of Scarlet Gilia (Ipomopsis aggregata) wave in the wind.

IpomAggr_flst_AntFl_6.19.17_1_crp_3x5_240The 1-2” red trumpet flowers attract hummingbirds, which are able to hover and extend their long tongues into the deep tube to lap up (not sip) nectar. A bird’s head may be doused in pollen on one visit. On the next stop, it is poked with a sticky stigma that will collect the pollen to make seeds. Pinkish flowers later in the season attract long-tongued sphinx moths, which provide a similar pollination service.

Lupines are another common flower of both sage flats and mountain slopes.

Silky Lupine - Lupinus sericeus

Silky lupine (Lupinus sericeus), which grows in sunny locations, has hairs on the backside of the “banner” of the pea-like flowers and very hairy palmate leaves. The hairs help protect plants from intense sun and wind of open sites.

In the Pea or Legume Family, lupines can “fix nitrogen”, enabling plants to grow in poor soils. Bacteria are harbored in nodules formed by the roots. In return for the plant’s protection and some food, bacteria convert nitrogen (NH2) from the air (soil has air pockets) into a form that plants can use (NH3). For centuries, farmers have grown clovers and alfalfa—also legumes–to provide this same soil enriching function.

Some hillsides along the Ski Lake Trail or under aspens at Munger Mountain are dominated by spires of yellow Fernleaf Lousewort (Pedicularis bracteosa). PediBrac_flhab_TetPsS_7.3.18_1b_Q2_5x3_180sm.jpgThese laterally flattened, irregular flowers require bumblebees to pollinate them. Bumblebees are strong and smart enough to land on the lower lip of the flowers and push and prod their way into the throat to find nectar. In so doing, the bee gets a bunch of pollen on its body. At another flower, it distributes pollen to the stigma protruding from the top of the upper lip.

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Flowers of louseworts are designed to fit specialized pollinators. Here you can see the stigma poised to tap pollen off the back of a visiting bee.

Fernleaf Louseworts are hemiparasites—they get extra nutrients and even chemical defenses from “host” plants. Roots of louseworts can attach to Arrowleaf Groundsel (see below) and Engelmann Spruce for these added benefits.

Another “free-loader” or hemi-parasite is Scarlet Indian Paintbrush (Castilleja miniata).   The plants attach underground “haustoria” to a variety of different species. Scarlet paintbrushes, and other paintbrush species, are blooming at different elevations in Jackson Hole.

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Cinquefoils (Potentilla spp.) appear pretty much everywhere. Research indicates that the yellow- to cream-colored, 5-petalled flowers of tall cinquefoils (Potentilla arguta/glandulosa) attract dozens of different types of pollinators, which is a good evolutionary trait for success. Different insects may or may not be abundant in different years.

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Sticky Cinquefoil (Potentilla arguta) is one of the most commonly seen species at this time. For precise ID, one counts up to 25 anthers, notes the roughly marked stigmas on smooth ovaries forming a slight cone in the center, and sees several flowers held tightly together on sticky stems.  In fact, taxonomists are lumping two look-alike species and now calling the genus Drymocallis. Definitely plant geek talk, you can ignore.

By being a generalist, cinquefoils are always likely to have some pollinators visit in any given year. Note: there are several different look-alike cinquefoils.

Stickseed (Hackelia micrantha) is abundant now at higher elevations, such as Teton Pass.

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The blue flowers of stickseed look like fragments of heavenly blue sky.

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However, the fruits will not be so delightful—they have devilish fruits with barbs that will attach to your socks. You will be their unwilling disperser to new lands in a few weeks.

A few other tall meadow flowers are seen along Moose-Wilson Road and will soon bloom up higher:

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A mix of tall flowers along Moose-Wilson Road where there is plenty of moistsure and sun.

Tall Larkspurs (Delphinium occidentale) are unravelling their deep- to pale-blue stalks of flowers.

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Tall Larkspur has trumpet-shaped flowers that require bee pollinators to push deeply into them to receiive their rewards.

Several tall groundsels (Senecio spp.) will soon add bouquets of yellow blossoms. Typically, flower heads all have several yellow ray (petal-like) flowers surrounded by a pallisade fence of even-sized green bracts—often with black tips. The leaves are helpful identifiers to species:

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Butterweed Groundsel (Senecio serra) has oblong serrated or toothed leaves. Plants grow to 4-6’ tall.

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Thickleaf Groundsel (Senecio crassulus) has somewhat fleshy or succulent, smooth leaves that clasp the stem. Plants  are around 12’ tall.

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Arrowleaf Groundsel (Senecio triangularis) are found in wet areas—seeps and stream edges. Leaves are distinctly arrow-shaped and sharply toothed. They grow to 2-3’ tall.

Can you guess what this is?

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A flower bud of Cow Parsnip (Heracleum spondylium)! Cow Parsnip has the largest flower cluster (umbel) of any member of the Parsley Family in the west.

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Cow Parsnip  (Heracleum spondylium) grows in moist meadows and ravines where there is enough water to supply the very large leaves on 4-6’ plants.

Of course there are many more flowers to see.  However, this “botanist’s dozen” is a good beginning to your explorations. Soon we will add postings for flowers growing in the forest, wetlands, and in just plain odd places.

Have fun!

Frances Clark, Teton Plants

P.S.  We try very hard to be accurate.  If you see an error, please let us know so that we can correct our mistakes at tetonplants@gmail.org .  Thank you!

Spring Emerging – April 2018

Our first flowers are finally revealing themselves as the snow melts along road verges, fields, sage flats, and open forests on the valley floor.

Many early wildflowers are “spring ephemerals”: they flower before there is competition for light by larger plants and then disappear, leaves and all, within a few weeks. They have adapted to this niche of opportunity.  Often just a few inches high, they are best observed on one’s belly – belly botany.

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A few sturdy woody plants are also blooming. At this time of year they count on wind for pollination, as insects are few. We often overlook their flowers because they don’t have showy petals: wind cannot see.TNP16_MWRd_AlnusSalix_Spr_1_5x3_180

Spring ephemerals emerge from underground storage units: tubers, bulbs, and rhizomes. Stored starch fuels new shoots to stretch above ground into the light where they can then form leaves for photosynthesis, making new food. They will quickly flower and then store fresh starch reserves underground for the next year. The leaves disappear from the surface—leaving only fruits to release seeds.

The growth pattern of our wild spring ephemerals is similar to our cultivated bulb plants, such as snowdrops, crocus, and daffodils whose foliage will fade by the end of spring. If you let them die back naturally in your garden instead of “tidying them up”, the leaves will make enough food to form new bulbs for a show next year.

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The flowers of Turkey PeasOrogenia linearifolia – are tiny: 10-12 or more blossoms will fit on your thumbnail. The plants are barely an inch or two high and hard to detect among old twigs, leaves, and stones. The name Turkey Pea likely comes from their tiny bulbs.

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Turkey peas are miniature members of the Parsley Family or umbellifers. The two parted stigma is maroon and surrounded by 5 white petals and maroon anthers. We observed flies and less frequently honey bees pollinating them.

Utah and Sage Buttercups are spreading their bright yellow petals–they gleam! Both species look very much the same; however,

Utah Buttercup – R. jovis has 3-parted leaves and fleshy, tuberous roots. So far I have seen these frequently under cottownwoods and in rough fields.

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Sage buttercup – R. glaberrimus – tends to have undivided leaves at least the first ones at the base.  Stem leaves may lobed.  The roots are cylindrical, not pudgy. As the name implies, it is more often found in sagebrush habitats.

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Honey bees are a pollinator to this Utah Buttercup.  Note Turkey peas in lower right. (Photo by Mary Lohuis 4.20.18.)

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Bending down low you can catch a whiff of their sweet fragrance 1-2’ off the ground. The sun warms the soil and wafts the scent to low-cruising pollinators—flies, bees, honeybees. They pick up the scent, then the color. The slight change in color in the inner part of petals is a change in the UV reflectance of “bee yellow”: the inner part is a contrasting bull’s-eye to the pollinator.

Spring BeautyClaytonia lanceolata – is beginning to appear. Two opposite leaves expand and 1 to several flowers will slowly stand up in between.   White to pinkish peals are striped pink, drawing in pollinators to open saucers of flowers serving nectar.

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YellowbellsFritillaria pudica – have been sighted! The 6 yellow tepals (3 sepals and 3 petals) dangle down forming a bell. Pollinators key into the changing of color at the base of the flower: green then reddish—indicating different stages of fertility.

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By all appearances Steer’s-heads – Dicentra uniflora – are the quintessential western spring flower with their distinctly bovine design.  The flowers serve to attract bees that can navigate the complex flowers to reach the nectar reward at the base. Bluish leaves are divided several times into rounded lobes and are toxic. Dicentra seeds are dispersed by ants.

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The Bleeding Heart (Fumatory) Family includes our ornamental bleeding hearts – Dicentra spectabilis – and our local species Golden Corydalis – Corydalis aurea – which can be seen along Game Creek in late April.

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Woody plants – With the lengthening of the days, buds begin to exude a hormone, auxin, which then spreads down woody stems stimulating cell division of the cambium—the stem tissue that encircles the stem just below the bark.  New vessels (xylem) enable water and nutrients from the roots to reach newly expanding shoots and flowers.

Typically, early spring shrubs and trees are wind pollinated. There is plenty of wind and relatively few insects about. Plants colonize open areas where there is little interference by leaves or trunks for pollen to blow from male to female flowers of the same species.

Most flowers are either male or female and come out at slightly different times or are on separate plants altogether to assure cross-fertilization—a mixing of genotypes.

AldersAlnus incana – are dangling their 2-3” long male catkins over the wetlands along Moose-Wilson Road and elsewhere. If you can get up close without getting your feet wet, you can look for the deep-maroon 1/4″ female catkins nearby on the same branch. They have scarlet stigmas which capture the pollen. You can also find last year’s tough 1” woody female “cones”. Male catkins wither away after they have released their pollen.

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AspensPopulus tremuloides – Fuzzy, silvery catkins are emerging on some trees but not others, depending on the clone. Male and female flowers are on separate trees and separate clones.  Below female catkins extend their maroon stigmas to catch the wind-dispersed pollen.

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Closely related to aspens, Cottonwood – Populus spp. – buds are bursting. Cottonwoods also have male and female plants. One can smell the distinctive odor of the balsam “oil”. The oil is popularly used as a salve and for aromatherapy.  In the photo below, male catkins are just emerging–note red anthers, also the sticky, fragrant sap on the bud scales.

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Pussy WillowsSalix spp. – Since childhood, many of us have loved pussy willows for their silky soft catkins. There are dozens of different types of pussy willows in Jackson Hole, with catkins ranging in size from ½-3” and the leaves of different sizes, shapes and textures. ID to species is very difficult, but the genus Salix is easy to determine. Buds have one covering or scale. Watch as this cap is pushed off as the catkins expand.

While I used to think willows were wind pollinated, in fact many willows are insect pollinated. The tiny scales hidden in the silvery hairs of the upright catkins have nectar glands at their base. UV light and perhaps the shiny catkin hairs attract bees and flies to this reward.  Vistors then carry pollen to a separate female plant. Look for anthers in the males  catkins (shown below) and stigmas in the females to know which gender the shrub is.

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Yellow willow stems are obvious in spring. Carotenoids produce yellow and orange hues (the same pigment that colors our carrots!).  These pigments help trap certain wavelengths to aid photosysntheis while at the same time protecting cells from harmful rays.  Willows are taking advantage of the bright unshadowed light for a jump start to growth in spring.

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April is a wonderfully subtle time of year when a few blooms count for much pleasure. I hope you can venture outside and enjoy it!

Frances Clark, Wilson, WY

4.22.18

As always, we appreciate any corrections.  Please email tetonplants@gmail.com

 

 

Wally’s World and Aspen Groves – late June 2017

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Wally’s World and Poison Creek Trails which loop around the shoulder of Munger Mountain encourage explorations into dry meadows and aspen groves.  Wally’s World Trail runs along an open ridge overwhelmed by mulesears – Wyethia amplexifolia – and abundant fragrant Antelopebrush.  This trail has many of the same sagebrush habitat plants noted in the recent 6.23.17 post (please see).

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AntelopebrushPurshia tridentata – mingles with sagebrush, snowberry, and mulesears along the ridge of Wally’s World, Bridger-Teton National Forest, off Fall Creek Road, Wilson.

Mixed among the sagebrush with Sulphur Buckwheat, Scarlet Gilia, Ballhead Sandwort, and Hawksbeards, are two different red paintbrushes.  PaintbrushesCastilleja spp. – are tricky plants to identify because of their modified flower parts and also because they hybridize and vary in color, leaf shape, and growth habit even within a given species. Interestingly, they also are hemiparasites.

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Paintbrush flowers are hard to identify in part because they have complicated flowers.  A colorful bract (shown to right) is under each flower. Each flower has a colorful calyx tube which is lobed. It surrounds the usually greenish galea = tube of fused petals. The stigma and anthers are protected inside. Here the stigma sticks out from the green and red galea.  If you are not in the national park and have plenty around, pick apart — dissect — a single flower and see what is what.

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One of the red paintbrushes Northwestern PaintbrushCastilleja angustifolia (C. Chromosa) – and Big SagebrushArtemesia tridentata var. vaseyana – mix together.  There is more to this view than meets the eye….

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Many paintbrushes are hemiparasites on a variety of hosts: grasses, legumes, and also sagebrush.  Northwestern PaintbrushCastilleja angustifolia  (C. chromosa) – attach “houstonia” to the roots of sagebrush and draw upon its water, nutrients, and some carbon to help the paintbrush grow more robustly.  This hemiparisite doesn’t depend entirely on the host: the chlorophyll in the leaves enables the plant to photosynthesize sugars (which include carbon).

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The state flower Wyoming PaintbrushCastilleja linariifolia – is a hemiparasite on sagebrush as well. In this paintbrush, the bracts are divided into linear lobes, the calyx is orange, toothed, with a long slit down the front where the elongate green galea leans out. This reddish tubular flower is pollinated by hummingbirds.

 

Aspen Forests are rich places to botanize.  BTNF_Mung_Aspen_6.22.17_6_Q2_5x3_200With light shade, greater moister, and more nutrients recycled from the deciduous aspen leaves, wildflowers are able to flourish at this time of year.

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The 3- to 4-foot Fernleaf LovageLigusticum filicinum – is beginning to flower. Note the large delicate leaves and umbels of tiny white flowers.

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In the same family – the Parsely or Apiaceae Family, SweetrootOsmorhiza occidentale – has tiny greenish yellow flowers in umbels and also large divided leaves.

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Sweetroot is just beginning to form its tangy fruit.  It is in the same family as many herbs we use: parsely, caraway, dill, fennel, etc.  While most are safe to taste, a few species in this family are highly toxic, such as poison hemlock.

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Also in the light shade of aspen groves or in open moist hillsides, are the yellow spires of Bracted LousewortPedicularis bracteosa.  These plants are also hemiparasites, operating like the paintbrushes, obtaining nutrients from surrounding plants.

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But perhaps equally interesting is its pollination.  Louseworts have very specific pollinators: bees that can just fit into the flower opening.  As they push inside for nectar, bees get a dusting of pollen on their bodies.  One pair of bee legs is designed for combing off pollen and stashing pollen grains in “buckets” on the hind pair of legs.  However, bee’s can’t reach the pollen lodged between their head and thorax.  When a bee visits the next flower, a perfectly shaped protruding stigma taps into that crevice and bingo the pollen gets stuck to the sticky stigma. The plant wins and seeds can now form!

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A much more unusual species, Tall LousewortPedicularis procera – can be seen on Munger Mountain.  The flowers are bigger and reddish, and the fern-like leaves are more frilly than the more common Bracted Lousewart.

Different species of lupines, paintbrushes, and even geranium are found in shadier woodland vs. open sage-covered habitats.

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Silvery LupineLupinus argenteus – is found most frequently in forests–aspen or pines.

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Silvery Lupine differs from the Silkly lupine, which is found in sageflats, by having many fewer silky hairs.  The banner–the top petal which folds backwards–is smooth, not hairy.  The flowers are slightly smaller and the leaves greener.   

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One-foot or more tall Scarlet PaintbrushCastilleja miniata – has bright red, unlobed bracts beneath the red flowers.  Also, the leaves are  usually not deeply lobed as are the two red paintbrushes observed in sunny, dry locations. 

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Sticky GeraniumGeranium viscosissimum – is abundant right now both on sageflats and in woodlands.  Note the nectar guides — the deep pink lines leading into the center of the flowers.  If you look closely with a handlens you can also see (and feel) the many sticky hairs on most parts of the plants.  The hairs happen to be clear.

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Richardson’s GeraniumGeranium richardsonii – is white and grows in moist sites.  Its hairs are different!

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The sticky hairs of Richardson’s Geranium are purple-tipped. Think of a gooey forest of defense against tiny invaders.

Several members of the Buttercup Family (Ranunculaceae) do well in the shade.  Many look very different and have very singular pollination behaviors from one another. Families are like that.

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Colorado ColumbineAquilegia coerulea – is very showy and a favorite to find. Elegant flowers have 5 flaring white petal-like sepals, 5 tubular petals that form 2”-long spurs trailing out the back, and many yellow anthers. Leaves are delicately dissected into 9 parts. These 6-24” plants are pollinated by moths and hummingbirds which have mouth parts that can reach the nectar way back in the flower “spurs”.

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In contrast to columbine, the flowers of MeadowrueThalictrum occidentale – are very inconspicuous.  It has male and female plants. Wind blows the pollen from the male anthers to the female stigmas on a nearby plant. This is a male plant. 

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Female flowers of Western MeadowrueThalictrum occidentale. The pink stigmas stretch to catch pollen grains.  As the wind cannot “see” there is no need for showy petals and such to attract pollinating birds or insects. Also, petals would get in the way of the wind.

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Fruits of Western Meadowrue are already forming.  There are actually two possible meadowrue species T. fenderi and T. occidentale.  The size and shape of the fruits help to distinguish the two species.  This observer does not swear to the species ID as T. occidentale.

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Yet another buttercup relative literally hangs out in the bushes.  Western ClematisClematis occidentalis – is a vine which curls its tendrils around branches to support itself.  The flowers nod downwards.  Often it is hard for us to see more than the back of the blue flowers; however, low flying pollinators look up into the blue to see hundreds of gleaming white stamens – a treasure trove of pollen. Heaven. 

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This cousin has many small white flowers whose numerous stamens (not petals) form a starry effect.  Placed together, the flowers create an appealing show to small pollinators.  

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While delicate in appearance, the flowers will produce ruby red berries that are poisonous–they were  used to make poison arrows.  Hence the name Red BaneberryActea rubra. Bane in a plant’s name indicates poisonous.  “Bane” means misery, misfortune, pain in the English language. 

After looking at the photos, can you see any family resemeblance among the buttecup relatives?  Hint: Look at leaves, look at stamens….(answer below)

Three additional flowers to look for in the forest at this time.

One is pretty obvious–reddish stems and spotted flowers:

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Spotted CoralrootCorallorhiza maculata – is growing along trail edges in aspen groves.  Coralroots do not have chlorophyll for photosynthesis and instead are connected to mychorrizal fungi which transport water, nutrients, and carbon from host trees.  There are several species of coralroot in Teton County.  This reddish species has white flowers spotted with red. Each flower has two tiny lobes on either side of the lowest petal (lip) and a bump (spur) under the throat . 

Two are tiny, easily overlooked or even stepped upon!

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These merry flowers are barely 1/4″ wide and decorate slender 12″ stems.  With the light behind them, Small-flowered Mitreworts – Mitella stauropetala – look like magical fairy wands.  

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Northwestern Twayblade – formerly Listera caurina, now Neottia  banksiana – is a 4-12″ orchid found in conifer forests (vs. aspen groves).  We came upon a few hidden clusters today.  It is found in Wyoming only in the Teton County.

There is no information on the pollination of Northwestern Twayblades–Listera caurina. However, one of its close relatives Listera cordata, which is also found in Teton County, has an amazing pollination mechanism that may be similar to this species. It is detailed below to encourge us to look for both species and do a close comparision,

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  Pictured is Northwestern Twayblade. The story below is about its relative Heart-leaved Twayblade.  First it is important to know that in orchids, the miniscule pollen grains are held in wads called pollinia…. Also the the anthers and pistil are fused into a single column or rostellum….  Attracted by the fetid odor and a bit of nectar, a small pollinator, such as  fungus gnat, comes to the flower. Rooting around the lower lip for nectar, it triggers one of three pressure-sensitive hairs on the arching upper part of the flower–the rostellum–above it.  At this point, the rostellum “releases a droplet of glue and the pollinia are instantly released to fall on the glue.”  Off the gnat goes with the pollinia stuck to its body.  Quickly, the tip of the rostellum unfolds to shield the underlying stigma to prevent self-fertilization in case the gnat returns. The next day, the wide stigma is uncovered and it arches up, receptive to the next gnat carrying pollinia from a different plant.  As the gnat crawls under the rostellum for nectar, the pollinia sticks to the stigma and hundreds of pollen grains germinate to form hundreds of dust like seeds. Thus begins the opportunity for new generations.  (Based on research of pollination of Listera cordata in California – Ackerman and Mesler, Am. Journal of Botanty, 1979)

Observing any twayblades in Jackson Hole, very carefully try to determine 1. odor, 2. presence of trigger hairs, 3. is the stigma exposed or covered? This could help determine if the pollination mechanisms within these two species are at all similar.

In any case, there are many more flowers to see!  Keep on botanizing!

Frances Clark, Wilson, WY – June 28, 2017

P.S. Many Buttercup Family relatives typically have deeply divided to compound leaves; highly modified sepals and petals or sometimes none; and many, many anthers. Delphinium/larkspur, monkshood, and sugar bowls are all related, as well.